Jonas  Rubenson 

photo of Jonas Rubenson

Associate Professor

Contact Information

29B Recreation Building
The Pennsylvania State University
University Park PA 16802


(fax) 814-863-4755

Research Interests

Fundamental principles underlying joint and muscle function during locomotion. We integrate experimental and modeling approaches to study gait and skeletal muscle function during locomotion in both health and disease/impairment. In particular, the relationship between joint and muscle mechanics and metabolic energetics and mechanisms underlying locomotor adaptation and optimization.


Ph.D., 2005, Biomechanics, The University of Western Australia


  • Mechanics and energetics of locomotion
  • In vivo skeletal muscle function
  • Musculoskeletal structure-function

Selected Publications

Pires, N.J., Lay, B. and Rubenson, J. (2014). Joint-level mechanical triggers for the walk-to-run transition in humans. J. Exp. Biol. 217: 3519-27.

Panizzolo, F.A., Maiorana, A.J., Naylor, L., Lloyd, D.G., Green, D.J. and Rubenson, J. (2014). Gait analysis in chronic heart failure points to the calf as the source of impaired walking capacity. J. Biomech. 47: 3719-25.

Stearne, S.M., Alderson, J.A., Green, B., Donnely, C.J. and Rubenson, J. (2014). Joint mechanics in rearfoot vs. forefoot running: implications of switching technique. Med. Sci. Sport. Exerc. 46:1578-87.

Rubenson, J., Pires, N.J., Loi, H.O., Pinniger, G.J., and Shannon, D. (2012). On the ascent: the soleus operating length is conserved to the ascending limb of the force length curve across gait mechanics in humans. J. Exp. Biol. 215: 3539 - 51.

Rubenson, J., Heliams, B.D., Besier, T.F.,Lloyd, D.A., and Fournier, P.A. (2011) Adaptations for economical running: the effect of bipedal limb structure on 3-D joint mechanics. Journal of the Royal Society, Interface. 8: 740-55.

Umberger, B.R. and Rubenson, J. (2011). Understanding muscle energetics in locomotion: New modeling and experimental approaches Exerc. Sport Sci. Rev. 39: 59-67.

Strategic Themes

  • Domains of Health and Behavior